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In males, nicotine affects the testes where prolonged exposure changes the morphology of spermatogenic cells, reduces sperm production, decreased steroidogenesis and inhibits spermatogenesis resulting in decreased sperm counts, poor sperm motility and increased levels of DNA damage [20, 21]

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Studies have shown that long non-coding RNA (lncRNA), microRNA (miRNA) and DNA methylation, through various mechanisms to regulate the transcription of genes and proteins, and activate a series of molecular signal pathways to participate in the process of apoptosis [44,45,46](Table 1)